aCore Center for Molecular Morphology, Section for Stereology and Microscopy, Aarhus University, Aarhus, Denmark

bDepartment of Pathology, Aarhus University Hospital, Aarhus, Denmark

cDepartment of Veterinary Clinical Sciences, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

dDanish Pain Research Center, Department of Clinical Medicine, Aarhus University, Aarhus, Denmark

eDepartment of Drug Design and Pharmacology, Faculty of Health and Medical Sciences, University of Copenhagen, Copenhagen, Denmark

*Corresponding author. Address: Department of Clinical Medicine, Core Center for Molecular Morphology, Section for Stereology and Microscopy, Aarhus University, Palle Juul Jensens Boulevard 99, C112, Level 1, DK-8200 Aarhus N, Denmark. Tel.: +45 2362 0153. E-mail address: [email protected] (D. Mortensen).

Sponsorships or competing interests that may be relevant to content are disclosed at the end of this article.

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D. Mortensen and M. S. Thoefner contributed equally to this work as co-first authors.

M. Berendt and J. R. Nyengaard contributed equally to this work as co-last authors.