aCentre for Neuroimaging Sciences, Institute of Psychiatry, Psychology and Neuroscience, King's College London, London, United Kingdom

bPain Research, Department of Surgery and Cancer, Faculty of Medicine, Imperial College London, London, United Kingdom

cWolfson Centre for Age Related Diseases, Institute of Psychiatry, Psychology and Neuroscience, King's College London, London, United Kingdom

dDepartment of Clinical and Biomedical Sciences, Faculty of Health and Life Sciences, University of Exeter, Exeter, United Kingdom

*Corresponding author. Address: Department of Clinical and Biomedical Sciences, Faculty of Health and Life Sciences, University of Exeter, Barrack Rd, Exeter, Devon EX2 5DW, United Kingdom. Tel.: +441392 72 8257. E-mail address: [email protected] (S. W. Hughes).

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