This work was supported by Shandong BellaGen Biotechnology Co., Ltd., and by grants from the National Natural Science Foundation of China (32188102 to J.-K.Z. and 32270050 to Ning J.), the Guangdong and Shenzhen Natural Science Foundation (2024A1515010541 and JCYJ20220530114409022 to Ning J.), the Guangdong Provincial Science and Technology Innovation Council Grant (2017B030301018). Ning J. is an investigator of Southern University of Science and Technology (SUSTech) Institute for Biological Electron Microscopy. We thank the staff at SUSTech Cryo-EM Center for assistance with data collection on the SUSTech Titan KRIOS cryo-electron microscope.