This work was supported by the National Natural Science Foundation (NSF) of China (82188101 and 32171236 to C.L.; 92353302 and 32170683 to D.L.; 81925012, 92049301 to B.L.), the Science and Technology Commission of Shanghai Municipality (STCSM) (22JC1410400 to C.L.), the Shanghai Pilot Program for Basic Research — Chinese Academy of Science, Shanghai Branch (JCYJ-SHFY-2022-005 to C.L.), the CAS Project for Young Scientists in Basic Research (YSBR-095 to C.L.), Shanghai Basic Research Pioneer Project to C.L, the Innovation Program of Shanghai Municipal Education Commission (2021-01-07-00-07.E00074 to B.L.). We thank the Cryo-Electron Microscopy center at Interdisciplinary Research Center on Biology and Chemistry, Shanghai Institute of Organic Chemistry for help with data collection. We thank the staff members of the Large Protein Preparation System/Nuclear Magnetic Resonance System at the National Facility for Protein Science in Shanghai (NFPS), Shanghai Advanced Research Institute, Chinese Academy of Sciences for providing technical support and assistance in data collection and analysis.