This work was supported by the Ministry of Science and Technology of China (grant no. 2023YFB3609000), the National Natural Science Foundation of China (grant nos. 52233010, 52103245, 22021002 and 22305252), the CAS Project for Young Scientists in Basic Research (grant no. YSBR-053), the Beijing National Laboratory for Molecular Sciences (BNLMS-CXXM-202012), China Postdoctoral Science Foundation (grant no. 2023M733555), and Postdoctoral Fellowship Program of CPSF (grant no. GZB20230771).