These authors contributed equally: Liang Chen, Shun Zhang, Niannian Xue, Mengjia Hong, Xiaohui Zhang.

Shanghai Frontiers Science Center of Genome Editing and Cell Therapy, Shanghai Key Laboratory of Regulatory Biology, Institute of Biomedical Sciences and School of Life Sciences, East China Normal University, Shanghai, China

Liang Chen, Shun Zhang, Niannian Xue, Mengjia Hong, Xiaohui Zhang, Dan Zhang, Jing Yang, Sijia Bai, Yifan Huang, Changming Luan, Biyun Zhu, Gaomeng Ru, Hongyi Gao, Meizhen Liu, Mingyao Liu, Yiyun Cheng, Liren Wang, Gaojie Song & Dali Li

Peking-Tsinghua Center for Life Sciences, Peking University, Beijing, China

Haowei Meng, Hao Wu & Chengqi Yi

National Center for International Biotargeting Theranostics, Guangxi Key Laboratory of Biotargeting Theranostics, Collaborative Innovation Center for Targeting Tumor Theranostics, Guangxi Medical University, Guangxi, China

Liping Zhong & Yongxiang Zhao