We thank Lu Zhou for help with proofreading the manuscript. This work was supported by the National Natural Science Foundation of China (82388201 to JH, 32170751 to Z-HY), the Fundamental Research Funds for the Central Universities (226-2024-00015 to XW), the National Key R&D Program of China (2020YFA0803500 to JH), the CAMS Innovation Fund for Medical Science (2019-I2M-5-062 to JH), Fujian Province Central to Local Science and Technology Development Special Program (No. 2022L3079 to JH) and the Fu-Xia-Quan Zi-Chuang District Cooperation Program (No. 3502ZCQXT2022003 to JH).