†Yingxi Wu and Haijian Wu contributed equally.

Center for Neurodegeneration and Regeneration, Zilkha Neurogenetic Institute, Room: 241, 1501 San Pablo Street, 90033, Los Angeles, CA, USA

Yingxi Wu, Haijian Wu, Jianxiong Zeng, Brock Pluimer, Shirley Dong, Xiaochun Xie, Xinying Guo, Tenghuan Ge, Xinyan Liang, Sudi Feng, Youzhen Yan, Naomi Sta Maria & Zhen Zhao

Department of Physiology and Biophysics, Keck School of Medicine, University of Southern California, 90033, Los Angeles, CA, USA

Yingxi Wu, Haijian Wu, Jianxiong Zeng, Brock Pluimer, Shirley Dong, Xiaochun Xie, Xinying Guo, Tenghuan Ge, Xinyan Liang, Sudi Feng, Youzhen Yan, Naomi Sta Maria & Zhen Zhao

Department of Neurosurgery, Second Affiliated Hospital, School of Medicine, Zhejiang University, 310009, Hangzhou, Zhejiang, China

Haijian Wu

Neuroscience Graduate Program, Keck School of Medicine, University of Southern California, 90033, Los Angeles, CA, USA

Brock Pluimer, Xinyan Liang & Zhen Zhao

Center for Craniofacial Molecular Biology, Herman Ostrow School of Dentistry, University of Southern California, 90033, Los Angeles, CA, USA

Jian‑Fu Chen

Lawrence D. Longo, MD Center for Perinatal Biology, Division of Pharmacology, Department of Basic Sciences, Loma Linda University School of Medicine, 92350, Loma Linda, CA, USA

Qingyi Ma

Brain Injury Research Center, Department of Neurosurgery, University of California, Los Angeles, 90095, Los Angeles, CA, USA

Fernando Gomez‑Pinilla