1Clinic for Transplantation Immunology and Nephrology, University Hospital Basel, Basel, Switzerland

2Department of Pathology, University Hospital Basel, Basel, Switzerland

3Molecular Immune Regulation, Department of Biomedicine, University of Basel, Basel, Switzerland

4HLA-Diagnostics and Immunogenetics, Department of Laboratory Medicine, University Hospital Basel, Basel, Switzerland

aCorresponding Author: Stefan Schaub, MD, Clinic for Transplantation Immunology and Nephrology, University Hospital Basel, Petersgraben 4, 4031 Basel, Switzerland, Tel: 0041 61 265 45 33, Fax: 0041 61 265 24 10, e-mail: [email protected], ORCID: 0000-0002-9170-1341

*contributed equally

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