ainAmind Laboratory, College of Life Sciences, University of Leicester, Leicester

bDepartment of Neurology, Queen Elizabeth Hospital

cNIHR King's Clinical Research Facility, King's College London, , London, UK

dDepartment of Neurology, University of California, Los Angeles, CA

eDepartment of Neurology, The Johns Hopkins University School of Medicine

fDepartment of Neuroscience, The Johns Hopkins University School of Medicine

gDepartment of Otolaryngology-Head and Neck Surgery, The Johns Hopkins University School of Medicine, Baltimore, Maryland, USA

Correspondence to Amir Kheradmand, MD, The Johns Hopkins Hospital, Department of Neurology, Vestibular and Ocular motor (VOR) Laboratory, Path 2-210, 600 N. Wolfe Street, Baltimore, MD 21287, USA. Tel: +1 410 955 3319; fax: +1 410 614 1746; e-mail: [email protected]