* Corresponding Author; email: [email protected]

1Department of Neuroscience and Pharmacology, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States

4Veterans Affairs Health Care System, Iowa City, Iowa, United States

5Department of Internal Medicine, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States

6Fraternal Order of Eagles Diabetes Research Center, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States

7Obesity Research and Education Initiative, University of Iowa Carver College of Medicine, Iowa City, Iowa, United States