These authors contributed equally: Xingyu Li, Peili Hou, Wenqing Ma, Xuefeng Wang.

Ruminant Diseases Research Center, College of Life Sciences, Shandong Normal University, Jinan, 250014, People’s Republic of China

Xingyu Li, Peili Hou, Wenqing Ma, Hongmei Wang, Zhangping Yu, Huasong Chang, Song Jin, Xue Wang, Wenqi Wang, Chunqing Xu, Xiaomei Ma & Hongbin He

Changchun Veterinary Research Institute, Chinese Academy of Agricultural Sciences, Changchun, 130122, People’s Republic of China

Xuefeng Wang, Tiecheng Wang, Wenqi Wang, Yudong Zhao, Yong Zhao & Yuwei Gao