*Address correspondence to: Rong Li (E-mail Address: [email protected]).

http://orcid.org/0000-0002-0540-6566

Department of Cell Biology, Center for Cell Dynamics, Johns Hopkins University School of Medicine, Baltimore, MD 21205, USA

Mechanobiology Institute, National University of Singapore, Singapore 117411, Singapore

Department of Chemical and Biomolecular Engineering, Whiting School of Engineering, Johns Hopkins University, Baltimore, MD 21218, USA

Department of Biological Sciences, National University of Singapore, Singapore, Singapore