These authors contributed equally: Pingping Zhu, Tiankun Lu, Jiayi Wu, Dongdong Fan.

CAS Key Laboratory of Infection and Immunity, CAS Center for Excellence in Biomacromolecules, Institute of Biophysics, Chinese Academy of Sciences, Beijing, China

Pingping Zhu, Tiankun Lu, Jiayi Wu, Benyu Liu, Hui Guo, Ying Du & Zusen Fan

School of Life Sciences, Zhengzhou University, Zhengzhou, Henan, China

Pingping Zhu

State Key Laboratory of Esophageal Cancer Prevention & Treatment, Zhengzhou University, Zhengzhou, Henan, China

Pingping Zhu

University of Chinese Academy of Sciences, Beijing, China

Tiankun Lu, Yong Tian & Zusen Fan

School of Medicine, Nankai University, 94 Weijin Road, Tianjin, China

Jiayi Wu

CAS Key Laboratory of RNA Biology; Institute of Biophysics, Chinese Academy of Sciences, Beijing, China

Dongdong Fan, Xiaoxiao Zhu & Yong Tian

Research Center of Basic Medicine, Academy of Medical Sciences, Zhengzhou University, Zhengzhou, Henan, China

Benyu Liu

State Key Laboratory of Membrane Biology, Institute of Zoology, Chinese Academy of Sciences, Beijing, China

Feng Liu