Available online 8 September 2022

The authors declare that they have no competing interests.

This work was supported by grants from the National Key R&D Program of China (2017YFA0105201, 2021YFA1101300, 2020YFA0112503), the Strategic Priority Research Program of the Chinese Academy of Science (XDA16010303), the National Natural Science Foundation of China (82030029, 81970882, 81900941, 82171144, 82192862, 81970884, 81771019.), the Natural Science Foundation from Jiangsu Province (BE2019711, BK20190121), the China Postdoctoral Science Foundation (2020M681555), the Outstanding Youth

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© 2022 Institute of Genetics and Developmental Biology, Chinese Academy of Sciences, and Genetics Society of China. Published by Elsevier Limited and Science Press.