The authors received funding from the Wellcome Trust (209192/Z/17/Z to C.S.), H2020 MSCA COFUND (663830-CU119 to C.S.), the Danish Foundation for Research in Neurology (to K.S.) and OPUS (grant 2017/27/B/HS6/00937 to K.S. and R.R.). In addition, the contributions of K.S. and R.R. to this article were based upon work from COST Action CA18106, supported by COST (European Cooperation in Science and Technology).